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Strawberry mottle virus, black raspberry virus and chocolate lily virus A are related to satsuma dwarf virus (SDV) in phylogenetic trees using the conserved Pro-Pol region (Figure 4.Secoviridae). Dioscorea mosaic associated virus, recently isolated from yam, is most closely related to chocolate lily virus A (Hayashi et al., 2016). These viruses also have a bipartite genome. However, the nature of their capsid protein(s) and their genomic organization are not known. For this reason, they are unassigned species in the family Secoviridae. Strawberry latent ringspot virus was formerly considered a sadwavirus because it has two capsid proteins (CP) and some distant relation with SDV in phylogenetic trees using the Pro-Pol sequence (Figure 4.Secoviridae). However, its genomic organization is more related to that of cheraviruses (with the exception of the number of CPs, Figure 3.Secoviridae) and it branches more closely with cheraviruses than with sadwaviruses in the phylogenetic trees using the Pro-Pol sequence (Figure 4.Secoviridae). For these reasons, it is not considered a sadwavirus anymore, and is now an unassigned species in the family Secoviridae.
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